Changing temperature regimes represent a severe threat to ecological communities, altering species growth, survival, and reproduction, which can result in increased extinction risks, changes to which species can coexist, and ultimately shifts in patterns of species diversity. However, to date, most studies of population and community responses to temperature regimes tend to examine either a handful of species or, alternatively, aggregated patterns like the total number of species (richness) in an ecosystem. The proposed research will improve understanding of how different temperature regimes alter ecological interactions at different scales. This CAREER project integrates research, education, and public engagement by creating a network – WyoCo Warming – of experimental sites with local communities experiencing both ambient and increased temperature conditions. Sites will be strategically placed across Colorado and Wyoming to engage local K-12 students, undergraduates, recreators, and land managers and will be include educational videos, K-12 lesson plans, and an educational website. WyoCo Warming sites will be monitored by undergraduate and graduate students as part of a new Community Ecology course. Creating a predictive framework for how changing temperature regimes will alter coexistence and richness patterns requires disentangling direct environmental effects from indirect effects of species interactions, both within the current community and in novel communities crea